1. Graham Proudlove's Cave Biology E-mail List Graham Proudlove's cave biology Email List. An email discussion group for cave biology exists. I hope that we can use it for respectable scientific debate about all aspects of cave biology. http://www.utexas.edu/depts/tnhc/.www/biospeleology/cavebiolist.htm

Graham Proudlove's Cave Biology E-mail List An email discussion group for cave biology exists. I hope that we can use it for respectable scientific debate about all aspects of cave biology. Please note the following details if you wish to take part in this list. To subscribe to the list send a message to: listserv@listserv.umist.ac.uk with the following one-line message: subscribe cave-biology To send a message to the list use the following address: cave-biology@listserv.umist.ac.uk To unsubscribe from the list send a message to: listserv@listserv.umist.ac.uk with the following one-line message: unsubscribe cave-biology Please don't send subscribe and unsubscribe messages to the list itself. Please let me know directly at g.proudlove@umist.ac.uk if you have any comments or suggestions about this list. [A note for UK readers - There is a specific email group for the UK. To subscribe to it, rather than the worldwide list (described above), substitute cave-biology-uk for cave-biology in the above examples.] Graham S. Proudlove

2. Biospeleology (cave Biology) Texas Memorial Museum's web site on biospeleology. Information on the biota of caves, karst, and groundwater. http://www.utexas.edu/depts/tnhc/.www/biospeleology/

Biospeleology The Biology of Caves, Karst, and Groundwater This page is sponsored by the Texas Memorial Museum, The University of Texas at Austin and the Missouri Department of Conservation Missouri Cave Life (pictorial guide) Explore Biospeleology (links) Handouts for cave managers (Elliott's files) NSS interview with Elliott A "cave salamander", Eurycea lucifuga Rafinesque, from Missouri. This species ranges across the eastern USA. This troglophilic amphibian prefers stream caves, primarily the twilight zone, but it also is found in damp habitats outside caves. Length 100-177 mm. This species has variable coloration, with dark or depigmented patches in some individuals. A few populations in Missouri have longer tails, similar to the long-tailed salamander. Both species love caves and frequently are found on walls or in crevices. Note the cirri on the upper lip, indicating that this is a male salamander. 18 May 2004 Comments to the Troglomeister: Bill.Elliott@mdc.mo.gov Missouri Department of Conservation

3. New York University | Cave Biology Research Group Main. Cave adapted animals are classic examples of regressive evolution, losing their eyes and pigment over time. Gains in other sensory systems compensate for the absence of vision. The bases for these developmental changes are genetic. The Mission. cave biology Facts. Research Programs http://www.nyu.edu/fas/cave

Main Cave adapted animals are classic examples of regressive evolution, losing their eyes and pigment over time. Gains in other sensory systems compensate for the absence of vision. The bases for these developmental changes are genetic. For these reasons, cave animals are an important resource to study the genetics and development of the visual system and other sensory modalities. Their study can help shed light on current problems in medicine and biology. Table of Contents Main The Mission Cave Biology Facts Research Programs Planned Expeditions ... Email to Webmaster

4. Biological Recorder A UK based organisation addressomg all aspects of cave biology. Papers, contacts, email discussion list, grid coordinates converter. http://www.bcra.org.uk/biology/

BCRA's Biological Recorder A Cave Biology special interest group does not exist within the British Cave Research Association , but we are pleased to announce that, from June 2000, we have resurrected the post of Biological Recorder . Our Recorder, Graham Proudlove, and his team of experts will be pleased to field your enquiries about cave biology. Graham's team hope to put together a web site for UK Cave Biology in the near future. In the meantime, Graham manages an e-mail discussion list and is in the process of assembling a bibliography of British subterranean biology. Cave Biology e-mail discussion list The Jefferson collection of cave fauna April 2003 Bibliography of British subterranean biology Jan 2004 A preliminary list of the obligate subterranean fauna of the British Isles Grid co-ordinates converter Many of the individual records contained within the "Biological Records" provide the 100km grid square details as a two digit number (e.g. 61/456789) rather than the more modern two letter code (e.g. SD/123456). Contact Details Graham Proudlove, Department of Zoology, The Manchester Museum, Manchester University, Manchester, M13 9PL. Tel (work) 0161 200 3111, Fax (work) 0161 200 3099, Tel (home) 01706 839752. E-mail

5. Illinois Natural History Survey, Biospeleology Web Site Links to other biospeleology sites cave biology ListServ and archives Biospeleologia del Piemonte, Atlante Fotografico Sistematico. cave biology Research Group ( New York University http://www.inhs.uiuc.edu/~sjtaylor/cave/biospeleol.html

ILDNR District Heritage Biologist Diane Tecic (L) and Tim Sickbert, an Illinois caver, (R) examine invertebrates on organic debris in a large Illinois cave. Photo (c) Steve Taylor Special Note: I n order to facilitate faster loading of this page, I have moved all of my cave critter pictures to a cave critter picture page [Top of Page] Karst Areas I n spite of Illinois' seemingly endless cornfields (formerly prairies), which occupy much of the central parts of the state, we are fortunate to have several hundred caves in four of Illinois' five karst regions along the southern, western, and northwestern borders of the state. Nearly all of our caves have developed in limestone bedrock. (Map courtesy of C. Pius Weibel and Samuel V. Panno, Illinois State Geological Survey - see literature section below for complete map citation [Top of Page] Terminology W ithin caves a diverse biota may be found, exhibiting varying degrees of adaptation to the subterranean environment. Accidental species, which fall, wander, or are washed into caves, do not linger long in this environment. These animals either return to the surface, or die in the caves - where they provide an important source of nutrients for the cave community. Trogloxenes occur commonly in caves, but must leave the cave at some point in their life cycle, typically for feeding. Species which occur in caves and can complete their entire life cycle there, but which are also found in similar habitats above ground, are referred to as

6. Utah Cave Biology NUTTY PUTTY. MISSION. CONSERVATION. GEOLOGY. MEETINGS. BIOLOGY. EDUCATION. IMAGES. VISITING CAVES Cave Biology. Underground Biology in Utah or a few cave locations; the pollution of a single http://www.caveutah.com/biology.htm

CAVE UTAH NUTTY PUTTY MISSION CONSERVATION GEOLOGY ... VISITING CAVES [Cave] Biology Underground Biology in Utah When asked about what lives in caves, most people respond 'bats'. However, many organisms live in caves, including beetles, spiders, crayfish, fish, salamanders, and more! Like bats, many creatures spend time in caves but must return to the surface in order to do certain things, such as finding food. Other creatures are able to spend their lives in caves, but can also be found on the surface. Yet others are found only in caves and nowhere else. Species that spend their entire lives in caves often exhibit specialized features as a result of many generations of subterranean living. Some of these features include the loss of pigment and eyes, the acquisition or exaggeration of non-visual sensory structures, and the elongation of the appendages. These cave-adapted species are often vulnerable to disturbance, causing population declines or even extinction, for several reasons. First, as a consequence of living a life underground where food supplies are often limited, many cave species have very low population numbers and reproduce infrequently Second, many species are found in only one or a few cave locations; the pollution of a single cave could cause some cave species to go extinct! In Utah, not much is known about what lives in the subterranean environment. Probably the most is known about bats; eighteen bat species have been found in Utah, with at least five of these species known to occur in caves and mines (

7. Cave Biology cave biology. There are several ways to look at cave biology and the ecology of Fisher Cave. The ecosystem approach is a useful beginning. http://www.missouri.edu/~jad4bc/biology.htm

Cave Biology There are several ways to look at cave biology and the ecology of Fisher Cave. The ecosystem approach is a useful beginning. In general, an ecosystem consists of biotic and abiotic elements interacting, while energy flows and nutrients cycle through the system. A cave is a low ordered systemin other words, it is relatively simple. First of all, caves lack the producer (or first trophic) level: there are no plants. Surface systems are usually diverse and complex, and thus stable. Caves, on the other hand, have low stability and are vulnerable to disturbance. As for trophic webs (that is, food chains), caves have a detritus food chain, not a grazing chain ( . The pathway from the base of the web is the breakdown of organic material, not plant-grazing. Energy is limited in caves: it comes second-hand from the surface, via a relatively small quantity of detritus. The conversion of energy from one level to the next is inefficient. Whereas most surface ecosystems convert energy at roughly a 10 percent rate (for example, from plant to herbivore to carnivore), in caves the conversion rate is about 2.5 percent. In a nutshell, a cave ecosystem is characterized by ecological simplicity, scarcity of energy (and thus food), and climatic stability.

8. Anchialine Caves And Cave Fauna - Biology Of Marine Caves marine cave biology, anchialine caves and cave animals, cave diving, marinebiospeleology, exploration and study of Yucatan and Bahamas caves. http://www.tamug.edu/cavebiology/

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9. Anchialine Caves And Cave Biology Peninsula of Mexico to carry out diving explorations and biological investigationsof the Ring of Cenotes, a 180 km diameter semicircle of water-filled caves. http://www.tamug.edu/cavebiology/index2.html

Bahamas Bermuda Yucatan Photo Galleries ... Site Map Thomas M. Iliffe e-mail: iliffe@cavebiology.com In March 2002, Tom Iliffe and his grad student Scott Webb were invited by film maker Wes Skiles to the Yucatan Peninsula of Mexico to carry out diving explorations and biological investigations of the Ring of Cenotes, a 180 km diameter semi-circle of water-filled caves. The Ring of Cenotes is the only surface expression of the 65 million year old meteorite impact crater which caused the extinction of the dinosaurs. The story of this expedition, "Watery Graves of the Maya", is recounted in the October 2003 issue of National Geographic Magazine and on the National Geographic website. To read about the expedition, see: http://magma.nationalgeographic.com/ngm/0310/feature4/index.html To view a photo of Tom Iliffe collecting an isopod from a Yucatan cave, see: http://magma.nationalgeographic.com/ngm/0310/feature4/zoom5.html To listen to Wes Skiles talk about the biological discoveries, click on Video 9 - Undiscovered Waters and Creatures: http://magma.nationalgeographic.com/ngm/0310/feature4/multimedia1.html

10. California Underground, Realms Educational materials and photographic tours on the formation and occurrence of different types of caves, cave biology, and cave conservation. http://www.museumca.org/caves/onli_realm.html

Our Caves and Subterranean Habitats Come explore the subterranean wonders of California. The diversity of the state's underground habitats is unsurpassed. We have sea caves, mud caves, lava tubes, talus caves, and wind caves, as well as the typical caves in limestone and marble. Click on this menu for a spectacular photo-tour of these realms. As you will see, the formation and occurrence of caves depends upon the interaction of rock types and climate within the context of the landscape. Where these factors combine favorably caves of a given type occur. Speleologists recognize many cave types and have proposed various schemes of categorizing them. For this web site we use a scheme of realms emphasizing how the caves were formed rather than rock type. © 1999 Oakland Museum of California Credits

11. Cave Biology Group cave biology Group. Our Recorder, Graham Proudlove, and his team of expertswill be pleased to field your enquiries about cave biology. http://www.bcra.org.uk/sig/biology.html

Cave Biology Group A Cave Biology 'special interest group' does not yet exist within the BCRA, but we are pleased to announce that, from June 2000, we have resurrected the post of Biological Recorder . Our Recorder, Graham Proudlove, and his team of experts will be pleased to field your enquiries about cave biology. Graham's team hope to put together a web site for UK Cave Biology in the near future. In the meantime, Graham manages an e-mail discussion list. BCRA's Biology Pages BCRA Home Page SIG Index Page Logos for Caving Groups David Gibson 29-Sep-00

12. Groundwater Biology Exhaustive collection of information about groundwater and cave biology. Checklist and bibliography for most taxa, distribution maps, images and useful links. Updated regularly. http://www.geocities.com/~mediaq/index1.html

13. New York University | Cave Biology Research Group | Facts cave biology Facts, cave biology Facts Cave vertebrates are excellent modelsto study the genetics of abnormal eye development and metabolic variation. http://www.nyu.edu/fas/cave/facts.htm

Cave Biology Facts Cave Biology Facts: Cave vertebrates are excellent models to study the genetics of abnormal eye development and metabolic variation. Retinal and lens defects in cave fishes are similar to those seen in humans and to those produced experimentally in the zebrafish development model. Karst systems are critical environmental resources; 40% of our drinking water passes through cave and karst systems. Cave fishes are sensitive indicators of environmental quality. Eye loss in cave fishes is the classic example of regressive evolution, an important phenomenon whose mechanism is still inadequately understood. Cave species have evolved independently, but converge towards a common theme. Their study brings the power of experimental replication to evolutionary biology and ecology, a power formerly restricted to laboratory science. Table of Contents Main The Mission Cave Biology Facts Research Programs Planned Expeditions Affiliations Advisory Board ... Email to Webmaster

14. Caving Canada Biospeleology Page Includes a directory of biospeleology researchers worldwide, Canadian cave biology directory, and many internet links. http://www.cancaver.ca/bio/

Caving Canada Biospeleology in Canada and on the WWW These pages on the Canadian Caving website are devoted to biospeleology: the scientific investigation of subterranean biology and the subterranean environment, which includes caves, groundwater and other underground habitats in Canada and throughout the world.These pages originated with Max Mosely (site curator 1996-1998) but are presently maintained by Internet Directory of Biospeleologists Bibliography of Canadian Biospeleology Local Site Search Guest Book ... The Canadian Cave and Karst Information Server This site created Sept '95, Updated Sunday, 25-Jan-2004 17:56:32 PST.

15. Cave Biology, Sequoia And Kings Canyon National Parks cave biology. People usually visit caves for the adventure of a newsubterranean world or to see beautiful cave formations. But, below http://www.nps.gov/seki/snrm/wildlife/cave_biology.htm

AIR GEOLOGY VEGETATION WATER ... HOME Wildlife Overview Bears Amphibians, Reptiles, and Fish Mammals ... Birds Cave Biology Wildlife Links and Lists Cave Biology People usually visit caves for the adventure of a new subterranean world or to see beautiful cave formations. But, below the feet and over the heads of visitors to caves are surprising and varied animals that make caves their home. Biologists divide cave-dwelling animals into three groups: Troglobites, which are restricted to caves; Troglophiles, who live in caves and on the surface; and Trogloxenes, who regularly visit caves but can not complete their life cycles in subterranean environments. Troglobites are often very specialized animals adapted to live in just one cave or a group of caves. Their adaptations may include no eyes or pigment, elongated appendages and enhanced tactile senses. These animals are very sensitive to change and disturbance. Troglobites or their food supply, can be trampled by caver visitors. Water pollution and surface developments may also poison cave environments. Bats are Trogloxenes, but they too are very sensitive to disturbance. They evolved to live in caves in complete darkness and silence. Unwitting cave visitors bring lights and make noises terrifying colonies of bats. Today a significant percentage of animals on the Endangered Species list is cave dwelling bats and invertebrates.

16. The Biospeleology In Romania About past and present cave biology research in Romania, including information about specific caves. http://www.uib.no/People/nglbn/bio.htm

The biospeleology in Romania General informations The first biospeleological researches in Romania were started by the Austrian and Hungarian zoologists, especially in Transylvania and Banat, while these regions were at that time under austro-hungarian occupation. They studied mainly the beetles, but neither the spiders, nor the myriapods were forgotten. In reality, the flourishing period for the Romanian biospeleology begins with the creation of the Speleological Institute in Cluj, in 1920. The triumvirate formed by Emil Racovita as manager, Rene Jeannel (from France) and Piere Alfred Chappuis (from Switzerland) as deputy managers started a research program covering the romanian territory and also other European and African countries. From biogeographical point o view the territory of the country can be divided in 5 biospeleological provices: the Oriental Carpathians as far as the valley of Olt, the Meridionals Carpathians between Olt, Timis and Cerna valleys, the Banat Mountains, the Apuseni Mountains and, Dobrogea. The karstificable surface represents 1.9 % of the romanian territory and its geographical distribution is unequal. This state is valuable also for the underground faunal distribution.

17. Jean Krejca, Vita Ralph Stone Award (National Speleological Society) spring 2001, for cave biologyresearch. Cave surveys and biological inventories were performed. http://www.inhs.uiuc.edu/~sjtaylor/cave/mexico/jkvita.html

JEAN K. KREJCA The University of Texas at Austin Integrative Biology PAT 140 1 University Station CO930 Austin, TX 78712-0253 creature@mail.utexas.edu Complete CV available as pdf file [20 kb]. Photo (c) Steve Taylor 1998 EDUCATION: Doctoral Candidate. University of Texas, Austin, doctoral research focuses on troglobites of Texas and Mexico B.S. in Zoology. Southern Illinois University with departmental honors for cave sculpin research, May 1995. RESEARCH INTERESTS: Cave biology, cave photography Texas field guide [note: all photographs in this link are © the photographer(s) ] Sequoia and King's Canyon National Parks cave biology inventory Aerial surveys using a powered paraglider Using aquifer organism genes as a dye tracer in karst Cave exploration and cartography projects Cave diving HONORS AND AWARDS: Mitchell Award (National Speleological Society) - August 1993, awarded for best scientific paper. Rope climbing awards (National Speleological Society) - August 1993, first place 120 and 30m 3 knot system. Cartography award (National Speleological Society) - August 1993, honorable mention for map of Clay Cave, Arkansas.

18. Onondaga Cave State Park - Cave Biology, Geology And History - Missouri State Pa cave biology, Geology and History. The History of Onondaga Cave Theland surrounding Onondaga Cave had been settled for some time http://www.mostateparks.com/onondaga/cavegeohist.htm

Cave Biology, Geology and History The History of Onondaga Cave The land surrounding Onondaga Cave had been settled for some time before the cave was discovered....... more information Geologic History of Onondaga Cave So what happened here at Onondaga? About a billion years ago, the Ozarks were a hotbed of volcanic activity centered about 45 miles to the southeast, in....... more information cave map Cave Geology A cave is a naturally occurring underground void, large enough to be examined in some way by man. In other words, a cave is a space, more than....... more information Speleothems Much less important to the geologist, but much more fascinating to the tourist are the speleothems, commonly called "cave formations."........ more information Cave Habitats You might get the impression that caves are beautiful, complex, fascinating and sterile. Sterile? Not on our living planet; caves are loaded with life........ more information Cave Animals The animals that are found in caves can be separated into five groups.......

19. The EnviroLink Network - Anchialine Caves And Cave Biology Anchialine Caves and cave biology. Diving explorations of anchialine (marine) caveshave led to the disovery of numerous animals, previously unknown to science. http://www.envirolink.org/resource.html?itemid=110302104197&catid=3

20. Correction For Anchialine Caves And Cave Biology Correction for Anchialine Caves and cave biology. Your Name YourEMail Your suggested changes for this resource Current Info http://www.envirolink.org/incorrect-info.html?itemid=110302104197&itemname=Anchi

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