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         Hartline Haldan Keffer:     more detail

81. Webvision: Ganglion Cell Physiology
In 1967 Ragnar Granit and H. keffer hartline shared the Nobel prize in physiologyand medicine for the first electrical recordings of light responses from
http://webvision.med.utah.edu/GCPHYS1.HTM
Visual Responses of Ganglion Cells Ralph Nelson [Overview] [History of Electrical Recordings] [ON and OFF Responses] [Receptive field] ... [References] 1. Overview. Ganglion cells are the final output neurons of the vertebrate retina. Ganglion cells collect information about the visual world from bipolar cells and amacrine cells (retinal interneurons). This information is in the form of chemical messages sensed by receptors on the ganglion cell membrane. Transmembrane receptors, in turn, transform the chemical messages into intracellular electrical signals. These are integrated within ganglion-cell dendrites and cell body, and 'digitized', probably in the initial segment of the ganglion-cell axon, into nerve spikes. Nerve spikes are a time-coded digital form of electrical signalling used to transmit nervous system information over long distances, in this case through the optic nerve and into brain visual centers. Ganglion cells are also the most complex information processing systems in the vertebrate retina. It is a general experimental truth that an organism as a whole cannot behaviorally respond to visual stimuli that are not also detectable by individual ganglion cells. Different cells become selectively tuned to detect surprisingly subtle 'features' of the visual scene, including color, size, and direction and speed of motion. These are called 'trigger features'. Even so signals detected by ganglion cells may not have a unique interpretation. Equivalent signals might result from an object changing brightness, changing shape, or moving. It is up to the brain to determine the most likely interpretation of detected events and, in the context of events detected by other ganglion cells, take appropriate action.

82. Ragnar Granit - By Yrsa Neuman/Gunnar Damström
equipment. At the same time British physiologist haldan keffer Hartlineconducted research at the University of Philadelphia. In
http://sfhs.eget.net/P_articles/Q_artiklar/Q_Granit.html
Ragnar Granit by Yrsa Neuman
At the University of Philadelphia Ragnar Granit had large research facilities at his disposal and received generous grants for acquisition of research equipment. At the same time British physiologist Haldan Keffer Hartline conducted research at the University of Philadelphia. In Pennsylvania Granit had the opportunity to experimantally test his theory on the nervous layer of the retina and its importance for seeing.
Ragnar Granit returned to Helsinki in 1934, and was shocked and saddened by the persecution of the Swedish language by the fennomans at the University of Helsinki. In his book he describes the hooligans hollering in the University aula and tarring Swedish language wall signs.
Granit wished to research whether the retina had separate receptors for the different components of solar light, in other words about the way the eye sees color. Granit reached conclusions similar to those of the classics of color vision theory the early English physicist Thomas Young and the German physicist and physiologist Hermann von Helmholz : that the retina has three color sensitive layers. It turned out that the eye entails a kind of nerve center that prepares the information for the brain and for the actual way there.
At the Carolinian Institute Granit focused his research on how the spinal marrow and the brain exert muscular control. He retired in 1967, the same year he shared the Nobel Price with Hartline and George Wald for his discoveries concerning the physiological and chemical foundations of vision. He passed away in 1991.

83. The Alfred B. Nobel Prize Winners: Physiology Or Medicine
United States United States. 1967, Ragnar Granit haldan keffer HartlineGeorge Wald, Sweden United States United States. 1968, Robert W
http://history1900s.about.com/library/misc/blnobelmed.htm
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Nobel Prize Winners for Physiology or Medicine
Chemistry Physics Literature Peace ... Economics Emil A. von Behring Germany Sir Ronald Ross Great Britain Niels R. Finsen Denmark Ivan P. Pavlov Russia Robert Koch Germany Camillo Golgi
Santiago Ramon y Cajal Italy
Spain Charles L. A. Laveran France Paul Ehrlich
Elie Metchnikoff Germany
France Emil Theodor Kocher Switzerland Albrecht Kossel Germany Allvar Gullstrand Sweden Alexis Carrel France Charles R. Richet France Robert Barany Austria Jules Bordet Belgium Schack A. S. Krogh Demmark Archibald V. Hill
Otto F. Meyerhof Great Britain
Germany Frederick G. Banting
John J. R. Macleod Canada
Scotland Willem Einthoven Netherlands Johannes A. G. Fibiger

84. Hartlines
hartline.com (Technology search firm); Town of hartline, WA. haldan Kefferhartline. Nobel biography www.nobel.se/laureates/medicine1967-2-bio.html.
http://www.pbrc.hawaii.edu/~danh/hartlines.html
Hartline Web Links
Other Hartline Links
Haldan Keffer Hartline

85. International: Italiano: Salute: Medicina: Medici_e_Ricercatori: Hartline,_Halda
Translate this page In tutta la Directory.
http://open-site.org/International/Italiano/Salute/Medicina/Medici_e_Ricercatori
Open Site The Open Encyclopedia Project Pagina Principale Aggiungi Contenuti Diventa Editore In tutta la Directory Solo in Medici_e_Ricercatori/Hartline,_Haldan_Keffer Top International Italiano Salute ... Medici e Ricercatori : Hartline, Haldan Keffer
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86. 1967Äêŵ±´¶ûÉúÀíѧ»òҽѧ½±»ñ½±Õß¹þÌØÀ¼´«ÂÔ
The summary for this Chinese (Simplified) page contains characters that cannot be correctly displayed in this language/character set.
http://www.med8th.com/nobel/winners/1967/Hartline-bg.htm
Haldan Keffer Hartline ÓÚ1903Äê12ÔÂ22ÈÕ³öÉúÔÚ±öϦ·¨ÄáÑǵIJ¼Â¬Ä·Ë¹±¤¡£ËûµÄ¸¸Ä¸¾ùÊÇÖÝÁ¢Ê¦·¶Ñ§Ð£(ÏÖÔڵIJ¼Â¬Ä·Ë¹±¤ÖÝÁ¢Ñ§Ôº)µÄ½Ìʦ¡£ËûÔÚÄÇÀï½ÓÊÜÁËÔçÄêµÄ½ÌÓý¡£ËûµÄ¸¸Ç×Daniel S.Hartline ÊÇÉúÎïѧ½ÌÊÚ£¬µ«¶ÔÌìÎÄ¡¢µØÀíµÈ¾ùÓй㷺µÄÐËȤ¡£¹þÌØÀ¼ÓÉÓÚÆ丸Ç×µÄÔµ¹Ê¿ªÊ¼¶Ô×ÔÈ»¿Æѧ·¢ÉúÐËȤ¡£
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1923ÄêÇËûµ½Ô¼º².»ôÆÕ½ð˹ҽѧԺ£¬ÔÚÉúÀíϵÖÐE.K.Marshall ºÍC.D.Snyder ¹ÄÀøËû¼ÌÐøÊÓ¾õ·½æµÄÑо¿£¬Snyder²©Ê¿ÈËûʹÓ×Ô¼ºµÄEinthovenÏÒÏßʽµçÁ÷¼Æ¡£¹þÌØÀ¼Ó´Ë»úÆ÷Ñо¿ÁËÍÜ¡¢È¥´óÄÔ¨ºÍ͵ÄÍøĤ¶¯×÷µçλ¡£Ëûѧ»áÁË´ÓÍêÕû¶¯ÎïÖлñµÍøĤµçͼ£¬²¢¼Ç¼ÏÂÇåÎú¿É±æµÄÈ˵ÄÍøĤ¶¯×÷µçλ¡£ËûÒ²ÓÍêÕûÀ¥³æ½øÐж¨Á¿Ñо¿
1927ÄêËû´ÓÔ¼º².»ôÆÕ½ð˹ҽѧԺµµ½Ò½Ñ§²©Ê¿ºó£¬¹ú¼ÒÑо¿Î¯Ô±»á½±Ñ§½ð(ҽѧ¿Æѧ)Ö§³ÖËûÑо¿ÊýѧºÍÎïÀíѧÒÔ¼ÓÇ¿Ëû½øÒ»²½Ñо¿ÉúÎïÎïÀíѧµÄ»ù´¡¡£ËûÔÚÔ¼º².»ôÆÕ½ð˹´óѧµÄÎïÀíϵ»¨·ÑÁËÁ½Äêʱ¼äÉϿβ¢ÒÔѧÉúµÈÉí·ÝÔÚA.H.PfundʵÑéÊÒ¹¤×÷¡£F.D.Murnaghan ÊÇËûµÄÊýѧÀÍʦ¡£1929Ä꣬Ëû´Ó±öϦ·¨ÄáÑÇ´óѧµµ½Eldridge Reeves Johnson ½±Ñ§½ð£¬¼ÌÐøËûÔÚÎïÀí·½æµÄÑо¿¡£Ëû»¨·ÑÁË°ëÄêʱ¼ä²Î¼ÓW.HeisenbergÔÚÀ´±ÈÎý´óѧ¿ªÉèµÄ½øÐް࣬ÓÁ½¸ö°ëÄêʱ¼ä²Î¼ÓĽÄáºÚ´óѧA.Somerfeld Ö÷³ÖµÄ½²ÑÝ¡£
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87. HighBeam Research: ELibrary Search: Results
Karl von Frisch, Ger.; Konrad Lorenz, Ger.Aus.; Nikolaas haldan KefferHartline, George Wald, both US 1966 Jacques Monod, all Fr.
http://www.highbeam.com/library/search.asp?FN=AO&refid=ency_refd&search_almanacs

88. The 125th Anniversary Of The Johns Hopkins University
Francis Peyton Rous, AB 1900; MD 1905, Medicine, 1966. haldan KefferHartline, MD 1927; Professor of Biophysics, 194954, Medicine, 1967.
http://www.jhu.edu/125th/links/nobel.html
Nobel Prize Winners with
Johns Hopkins Affiliations Name Affiliation Category Year Woodrow Wilson
Ph.D. 1886 (History) Peace James Franck Professor of Physics, 1935-38 Physics Nicholas Murray Butler Lecturer, 1890-91 Peace Thomas Hunt Morgan Ph.D. 1890 (Zoology); LL.D. 1915 Medicine George Richards Minot Assistant in Medicine, 1914-15 Medicine George Hoyt Whipple M.D. 1905; Associate Professor in Pathology, 1910-14 Medicine Harold Clayton Urey Associate in Chemistry, 1924-28 Chemistry Joseph Erlanger M.D. 1899; Assistant in Physiology, 1900-01; Instructor, 1901-03; Associate, 1903-04; Associate Professor, 1904-06; LL.D. 1947 Medicine Herbert Spencer Gasser M.D. 1915 Physiology Vincent du Vigneaud National Research Fellow, Pharmacology 1927-28 Chemistry Maria Goeppert-Mayer Assistant in Physics, 1930-32; Associate, 1932-36 Physics Francis Peyton Rous A.B. 1900; M.D. 1905 Medicine Haldan Keffer Hartline M.D. 1927; Professor of Biophysics, 1949-54 Medicine Lars Onsager Associate in Chemistry, 1927-28 Chemistry Simon Kuznets Professor of Political Economy, 1954-60 Economics Christian B. Anfinsen

89. Results Of Your Directory Search Of Brain Injury Agency Libraries
182. Visual Receptors and Retinal Interaction (alt1) Author(s) hartline, HaldanKeffer Located at ARBI Search Google for Visual Receptors and Retinal Intera
http://www.brainresource.ca/search_Directory.asp?q=psychology&p=19

90. Dorlands Medical Dictionary
American biologist, born 1906; cowinner, with Ragnar Arthur Granit and HaldanKeffer hartline, of the Nobel prize for medicine or physiology in 1967 for
http://www.mercksource.com/pp/us/cns/cns_hl_dorlands.jspzQzpgzEzzSzppdocszSzuszS

91. Nobel Laureates At Penn
1956 LL.D. 1976; Nobel Foundation information on this award. HaldanKeffer hartline, 1903 1983 Medicine, 1967. with George Wald and
http://www.archives.upenn.edu/histy/notables/awards/nobel.html
University Archives and Records Center
University of Pennsylvania Nobel Laureates
at the University of Pennsylvania
Awarded annually since 1901 by the Nobel Foundation , Stockholm.
Raymond Davis, Jr.
Physics, 2002
  • With Masatoshi Koshiba (University of Tokyo, Japan) and Riccardo Giannoni (Associated Universities Inc). Awarded in recognition of their groundbreaking research into the emission of neutrinos produced by nuclear fusion reactions in the center of the sun. The observation of these neutrinos demonstrated conclusively that the sun is powered by the fusion of hydrogen nuclei into helium nuclei. Davis joined Penn's faculty in 1985 after 37 years at Brookhaven Lab. Earlier this year, Davis received the 2001 National Medal of Science from President George W. Bush. Nobel Foundation information on this award.
Alan G. MacDiarmid

92. Warren Medal
No Award. 1945. Clark L. Hull. 1946. No Award. 1947. Wolfgang Kohler. 1948. HaldanKeffer hartline. 1949. Walter R. Miles. 1950. Curt P. Richter. 1951. Frank A. Beach.1952.
http://www.sepsych.org/warren_medal.htm
Howard Crosby Warren Medal
The Society of Experimental Psychologists awards the Howard Crosby Warren Medal annually for outstanding achievement in Experimental Psychology in the United States and Canada. Warren Medal Awardees, 1936 - present: Karl Spencer Lashley Elmer A. Culler Carlyle Jacobsen Ernest R. Hilgard Clarence H. Graham B. F. Skinner Stanley Smith Stevens No Award Clark L. Hull No Award Wolfgang Kohler Haldan Keffer Hartline Walter R. Miles Curt P. Richter Frank A. Beach James J. Gibson Kenneth W. Spence Neal E. Miller Georg von Bekesy Harry F. Harlow Lorrin A. Riggs Donald O. Hebb Harry Helson Carl Pfaffmann Carl I. Hovland James Olds William K. Estes Benton J. Underwood William C. Young Floyd Ratliff Eliot Stellar Richard L. Solomon Roger W. Sperry Daniel S. Lehrman George A. Miller Delos D. Wickens Leo J. Postman Wendell R. Garner Eleanor J. Gibson John Garcia Russell L. DeValois Abram Amsel Roger N. Shepard Endel Tulving Richard M. Held Eric R. Kandel Gordon H. Bower Hans Wallach Michael I. Posner Richard F. Thompson Anne M. Treisman

93. ÇÏƲ¶óÀÎ
The summary for this Korean page contains characters that cannot be correctly displayed in this language/character set.
http://preview.britannica.co.kr/spotlights/nobel/list/B24h2424a.html
Haldan Keffer Hartline ½°¢ÀÇ ½Å°æ»ý¸®ÇÐÀû ¸ÞÄ¿´ÏÁòÀ» ºÐ¼®ÇÑ °ø·Î·Î G.¿ùµå, R.±×¶ó´ÏÆ®¿Í ÇÔ²² 1967³â ³ëº§ »ý¸®ÇС¤ÀÇÇлóÀ» °øµ¿ ¼ö»óÇß´Ù. ÇÏƲ¶óÀÎÀº 1927³â º¼Æ¼¸ð¾îÀÇ Á¸½ºÈ©Å²½º´ëÇб³¿¡¼­ ÀÇ»çÀÚ°ÝÀ» ȹµæÇÑ ÈÄ, µ¿ ´ëÇб³ ºÎ¼Ó ±¹¸³¿¬±¸½ÉÀÇȸÀÇ Æç·Î·Î¼­ ¸Á¸·ÀÇ Àü±â»ý¸®Çп¡ °üÇØ ¿¬±¼¸£ ÁýÁßÇϱ⠽ÀÛÇß´Ù. ¿¤µå¸®Áö Á¸½¼ °´¿ø¿¬±¸ÇÐÀڷμ­ ¶óÀÌÇÁÄ¡È÷´ëÇб³¿Í ¹ÀÇî´ëÇб³¿¡¼­ ¿¬±¸ÇÑ ÈÄ 1949³â Á¸½ºÈ©Å²½º´ëÇб³ÀÇ »ý¹°¹°¸®Çаú ±³¼ö ¹× °úÀåÀÌ µÇ¾ú´Ù. 1953³â ½Å°æ»ý¸®ÇÐ ±³¼ö·Î¼­ ´º¿å ½ÀÇ ·ÏÆç·¯´ëÇб³ÀÇ ±³¼öÁø¿¡ ÇÕ·ùÇß´Ù. Limulus polyphemus

94. DICCIONARIO DE TERMINOLOGIA MEDICA

http://www.iqb.es/diccio/n/nobel07.htm
Premios Nobel
Von BEKESY, Georg (1899-1972). Estados Unidos CRICK, Francis Harry Compton WATSON, James Dewey (n.1928). Estados Unidos, WILKINS, Maurice Hugh por sus descubrimientos sobre la estructura del DNA ECCLES Sir John Carew (1903-1997) Australia, HODGKIN Alan Lloyd HUXLEY Andrew Fielding BLOCH Konrad (1912-2000) Estados Unidos; LYNEN, Feodor (1911-1979) Alemania JACOB François (n. 1920) Francia, LWOFF, André (1902-1944) Francia, MONOD, Jacques (1910-1976) Francia ROUS, Peyton (1879-1970) Estados Unidos, HUGGINS Charles Brenton (1901-1998) Estados Unidos GRANIT, Ragnar (1900-1991) Suecia, HARTLINE Haldan Keffer (1903-1983) Estados Unidos; WALD George HOLLEY Robert W. (1922-1993), Estados Unidos, KHORANA, Har Gobind (n 1922) Estados Unidos, NIRENBERG Marshall W. (n. 1927) Estados Unidos DELBRUCK, Max (1906-1981), Alemania; HERSEY, Alfred

95. Lexikon - 1967 Definition Erklärung Bedeutung
- Miguel Angel Asturias; Friedensnobelpreis - (nicht verliehen). Musik.
http://www.net-lexikon.de/1967.html

96. ¢Æ¢Æ¢Æ û¼Ò³â °úÇи¶´ç¿¡ ¿À½Å°ÍÀ» ȯ¿µÇÕ´Ï
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http://www.most.go.kr/most/Young_most/novel1.html
von Behring, Emil Adolf Ross, Sir Ronald Finsen, Niels Ryberg Pavlov, Ivan Petrovich Koch, Robert Golgi, Camillo Laveran, Charles Louis Alphonse Ehrlich, Paul Metchnikoff, Elie Kocher, Emil Theodor Kossel, Albrecht Gullstrand, Allvar Carrel, Alexis Richet, Charles Robert
Bordet, Jules Krogh, Schack August Steenberg
Hill, Sir Archibald Vivian Banting, Sir Frederich Grant Macleod, John James Richard Einthoven, Willem
Fibiger, Johannes Andreas Grib Wagner-Jauregg, Julius Nicolle, Charles Jules Henri Eijkman, Christiaan Hopkins, Sir Frederick Gowland Landsteiner, Karl Warburg, Otto Heinrich Sherrington, Sir Charles Scott Adrian, Lord-Edgar Douglas Morgan, Thomas Hunt Whipple, George Hoyt Minot, George Richards Murphy, William Parry Spemann, Hans Dale, Sir Henry Hallett Domagk, Gerhard
Dam, Henrik Carl Peter Doisy, Edward Adelbert ºñŸ¹Î K ¹ß°ß ºñŸ¹Î KÀÇ È­ÇÐÀû º»¼º ¹ß°ß Erlanger, Joseph Gasser, Herbert Spencer Fleming, Sir Alexander Chain, Sir Ernst Boris Florey, Lord(Howard Walter) Muller, Hermann Joseph X¼±¿¡ ÀÇÇÑ Àΰø µ¹¿¬º¯ÀÌ ¿¬±¸ Houssay, Bernardo Alberto

97. ƒm[ƒxƒ‹ˆãŠwÜ
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http://www.eps4.comlink.ne.jp/~simoyama/novelig-e.htm
ƒm[ƒxƒ‹ˆãŠwE¶—ŠwÜ ”N“x ˆãŠwE¶—ŠwÜ E.A.vonƒx[ƒŠƒ“ƒO ƒhƒCƒc Emil Adolf von Behring R.ƒƒX ƒCƒMƒŠƒX Ronald Ross N.ƒtƒBƒ“ƒZƒ“ Niels Ryberg Finsen I.P.ƒpƒuƒƒt ƒƒVƒA Ivan Petrovich Pavlov R.ƒRƒbƒz ƒhƒCƒc Robert Koch C.ƒSƒ‹ƒW ƒCƒ^ƒŠƒA Camillo Golgi S.ƒ‰ƒ‚ƒ“EƒCEƒJƒnƒ‹ ƒXƒyƒCƒ“ Santiago Ramon y Cajal C.L.A.ƒ‰ƒuƒ‰ƒ“ ƒtƒ‰ƒ“ƒX Charles Louis Alphonse Laveran P.ƒG[ƒ‹ƒŠƒq ƒhƒCƒc Paul Ehrlich E.ƒƒ`ƒjƒRƒt ƒƒVƒA Ilya Ilyich Mechnikov E.T.ƒRƒbƒqƒƒ[ ƒXƒCƒX Emil Theodor Kocher A.ƒRƒbƒZƒ‹ ƒhƒCƒc Albrecht Kossel A.ƒOƒ‹ƒXƒgƒ‰ƒ“ƒh ƒXƒEƒF[ƒfƒ“ Allvar Gullstrand A.ƒJƒŒƒ‹ ƒtƒ‰ƒ“ƒX Alexis Carrel C.R.ƒŠƒVƒF ƒtƒ‰ƒ“ƒX Charles Robert Richet R.ƒoƒ‰ƒj[ ƒI[ƒXƒgƒŠƒA Robert Barany ƒxƒ‹ƒM[ Robert Barany A.ƒNƒ[ Schack August Steenberg Krogh A.V.ƒqƒ‹ ƒCƒMƒŠƒX Archibald Vivian Hill ƒhƒCƒc Otto Fritz Meyerhof F.ƒoƒ“ƒeƒBƒ“ƒO ƒJƒiƒ_ Frederick Grant Banting ƒCƒMƒŠƒX John James Richard Macleod W.ƒAƒCƒ“ƒgƒz[ƒtƒFƒ“ ƒIƒ‰ƒ“ƒ_ Willem Einthoven J.ƒtƒB[ƒrƒK[ Johannes Andreas Grib Fibiger J.W.vonƒ„ƒEƒŒƒbƒO ƒI[ƒXƒgƒŠƒA Julius Wagner-Jauregg C.J.H.ƒjƒRƒ‹ ƒtƒ‰ƒ“ƒX Charles Jules Henri Nicolle F.G.ƒzƒvƒLƒ“ƒY

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